Thomas Tscheulin

T  (+30) 22510 36463
F  (+30) 22510 36423
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I was born in a small town in Germany right on the border with France and Switzerland. During my studies of Biology at the University of Freiburg (Germany) I spent two trimesters at the University of Reading (UK). This “short stint” in the UK should last almost a decade and so after finishing my degree in Freiburg I returned to the UK where I completed a PhD in Population Ecology at Imperial College, London and then went on to work at the Centre of Agri-Environmental  Research (CAER), University of Reading (UK) as a research assistant and post-doctoral research fellow. At CAER I was mainly involved in projects looking at ways to measure and enhance insect biodiversity in cultivated areas. 

Here at the University of the Aegean I am an Assistant Professor teaching the courses (or parts of) Environment & Ecology, Ecogeography, Biogeography, Environmental Impact Assessment, Areas for Conservation (postgraduate), Environmental Threats and Dangers (postgraduate), Landscape Ecology (currently not offered) and Environmental Geography.

My main research interest is to relate assessments of the abundance, diversity, functional structure and trophic interactions of invertebrate taxa, that are key in the provision of ecosystem services (e.g. pollination, nutrient cycling and biocontrol), to impacts of ecosystem disturbances such as agricultural practices, alien species invasion, climate change, wildfires, habitat loss and degradation, at a local, landscape and regional scale. Since my time at CAER (2004) I am interested in pollinators (mainly wild bees and Syrphidae) and, together with my colleagues here, study their ecology and distribution.

I like spending my free time with my wife and my son enjoying the great outdoors, camping, climbing, hiking, etc.


Lazarina, M., Devalez, J., Neokosmidis, L., Sgardelis, S.P., Kallimanis, A.S., Tscheulin, T., Tsalkatis, P., Kourtidou, M., Mizerakis, V., Nakas, G., Palaiologou, P., Kalabokidis, K., Vujic, A., Petanidou, T. 2019. Moderate fire severity is best for the diversity of most of the pollinator guilds in Mediterranean pine forests. Ecology e02615.

Kaloveloni, A., Tscheulin, T., Petanidou, T. 2018. Geography, climate, ecology: What is more important in determining bee diversity in the Aegean Archipelago? Journal of Biogeography 45 (12): 2690-2700.

Takkis, K., Tscheulin, T., Petanidou, T. 2018. Differential effects of climate warming on the nectar secretion of early-and late-flowering Mediterranean plants. Frontiers in Plant Science 9, 874.

Kantsa A, Raguso RA, Dyer ΑG, Olesen JM, Tscheulin T, Petanidou T. 2018. Disentangling the role of floral sensory stimuli in pollination networks. Nature Communications, 9:1041.

Petanidou T, Price MV, Waser NM, Kantsa A, Tscheulin T, Kariyat RR, Krigas N, Mescher MC, De Moraes CM, Bronstein JL. 2018. Pollination and reproduction of an invasive plant inside and outside its ancestral range. Acta Oecologica, 89: 11-20.

Petanidou, T., Kallimanis, A. S., Lazarina, M., Tscheulin, T., Devalez, J., Stefanaki, A., Hanlidou, E., Vujić, A., Kaloveloni, A. & Sgardelis, S. P. 2018. Climate drives plant–pollinator interactions even along small-scale climate gradients: the case of the Aegean. Plant Biology, 20, 176-183.

Gavalas V., Tscheulin T. 2017. Diverse marriage patterns in Imperial Germany. Journal of Family History 42(1):37-53.

Lazarina, M., Sgardelis, S.P., Tscheulin, T., Devalez, J., Mizerakis, V., Kallimanis, A.S, Papakonstantinou, S., Kyriazis, T., Petanidou, T. 2017. The effect of fire history in shaping diversity patterns of flower-visiting insects in post-fire Mediterranean pine forests. Biodiversity and Conservation 26(1): 115–131.

Neokosmidis L., ,Tscheulin, T., Devalez, J., Petanidou, T. 2016. Landscape spatial configuration is a key driver of wild bee demographics. Insect Science (in press).

Goras, G., Tananaki, C., Dimou, M., Tscheulin, T., Petanidou, T., Thrasyvoulou, A. 2016. Impact of honeybee (Apis mellifera L.) density on wild bee foraging behaviour. Journal of Apicultural Science 6(1): 49-61.

Lazarina, M., Sgardelis, S.P., Tscheulin, T.,  Kallimanis, A.S., Devalez, J., Petanidou, T. 2016. Bee response to fire regimes in Mediterranean pine forests: The role of nesting preference, trophic specialization, and body size. Basic and Applied Ecology 17: 308-320.

Lázaro A., Tscheulin T., Devalez J., Nakas G., Stefanaki A., Hanlidou E., Petanidou T. 2016. Moderation is best: effects of grazing intensity on pollination networks in Mediterranean communities. Ecological Applications 26(3): 796-807.

Lázaro, A., A. Chroni, T. Tscheulin, J. Devalez, C. Matsoukas, and T. Petanidou. 2016. Electromagnetic radiation of mobile telecommunication antennas affects the abundance and composition of wild pollinators. Journal of Insect Conservation: 20(2): 315-324.

Vujić, A., T. Petanidou, T. Tscheulin, P. Cardoso, S. Radenković, G. Ståhls, Ž. Baturan, G. Mijatović, S. Rojo, C. Pérez-Bañón, J. Devalez, A. Andrić, S. Jovičić, D. Krašić, Z. Markov, D. Radišić, and G. Tataris. 2016. Biogeographical patterns of the genus Merodon Meigen, 1803 (Diptera: Syrphidae) in islands of the eastern Mediterranean and adjacent mainland. Insect Conservation and Diversity 9: 181-191.

Lázaro A., Tscheulin T., Devalez J., Nakas G., Petanidou T. 2016. Effects of grazing intensity on flower cover, pollinator abundance and diversity, and pollination services. Ecological Entomology 41(4): 400-412.

Takkis, K., Tscheulin, T., Tsalkatis, P., Petanidou, T. 2015. Climate change reduces nectar secretion in two common Mediterranean plants. AoB Plants 7:plv111 pdf

Stefanaki, A., Kantsa, A., Tscheulin, T., Charitonidou, M & Petanidou, T. 2015. Lessons from Red Data Books: Plant vulnerability increases with floral complexity. PLOS ONE, 10(9): e0138414.

Kaloveloni, A., T. Tscheulin, A. Vujić, S. Radenković, and T. Petanidou. 2015. Winners and losers of climate change for the genus Merodon (Diptera: Syrphidae) across the Balkan Peninsula. Ecological Modelling 313:201-211. pdf

Carvalheiro, L. G., J. C. Biesmeijer, G. Benadi, J. Fründ, M. Stang, I. Bartomeus, C. N. Kaiser-Bunbury, M. Baude, S. I. F. Gomes, V. Merckx, K. C. R. Baldock, A. T. D. Bennett, R. Boada, R. Bommarco, R. Cartar, N. Chacoff, J. Dänhardt, L. V. Dicks, C. F. Dormann, J. Ekroos, K. S. Henson, A. Holzschuh, R. R. Junker, M. Lopezaraiza-Mikel, J. Memmott, A. Montero-Castaño, I. L. Nelson, T. Petanidou, E. F. Power, M. Rundlöf, H. G. Smith, J. C. Stout, K. Temitope, T. Tscharntke, T. Tscheulin, M. Vilà, and W. E. Kunin. 2014. The potential for indirect effects between co-flowering plants via shared pollinators depends on resource abundance, accessibility and relatedness. Ecology Letters 17:1389-1399. pdf

Nayak, K. G., S. P. M. Roberts, M. Garratt, T. D. Breeze, T. Tscheulin, J. Harrison-Cripps, I. N. Vogiatzakis, M. T. Stirpe, and S. G. Potts. 2015. Interactive effect of floral abundance and semi-natural habitats on pollinators in field beans (Vicia faba). Agriculture, Ecosystems and Environment 199:58–66. pdf

Bartomeus, I., Potts, S.G., Steffan-Dewenter, I., Vaissière, B.E., Woyciechowski, M.,Krewenka, Κ.Μ., Tscheulin, Τ., Roberts, S.P.M., Szentgyörgyi, H., Westphal, C., Bommarco, R. 2014. Contribution of insect pollinators to crop yield and quality varies with agricultural intensification. PeerJ 2:e328

Kantsa, A., T. Tscheulin, R. R. Junker, T. Petanidou, and S. Kokkini. 2013. Urban biodiversity hotspots wait to get discovered: The example of the city of Ioannina, NW Greece. Landscape and Urban Planning 120:129-137. pdf

Petanidou T., Ståhls G., Vujić A., Olesen J.M., Rojo S., Thrasyvoulou A., Sgardelis S., Kallimanis A.S., Kokkini S., Tscheulin T. 2013. Investigating Plant–Pollinator Relationships in the Aegean: the approaches of the project POL-AEGIS (The Pollinators of the Aegean Archipelago: Diversity and Threats). Journal of Apicultural Research, 52:106-117.

Tscheulin, T., and T. Petanidou. 2013. The presence of the invasive plant Solanum elaeagnifolium deters honeybees and increases pollen limitation in the native co-flowering species Glaucium flavum. Biological Invasions, 15:385-393. pdf

Nielsen, A., Dauber, J., Kunin, W.E., Lamborn, E., Jauker, B., Moora, M., Potts, S.G., Reitan, T., Roberts, S., Sõber, V., Settele, J., Steffan-Dewenter, I., Stout, J.C., Tscheulin, T., Vaitis, M., Vivarelli, D., Biesmeijer, J.C., Petanidou, T. 2012. Pollinator community responses to the spatial population structure of wild plants: A pan-European approach. Basic and Applied Ecology, 13:489-499. pdf

Tscheulin, T., Neokosmidis, L., Petanidou, T. and J. Settele. 2011. Influence of landscape context on the abundance and diversity of bees in Mediterranean olive groves. Bulletin of Entomological Research, 5:557-564. pdf

Nielsen, A., SteffanDewenter, I., Westphal, C., Messinger, O., Potts, S.G., Roberts, S.P.M., Settele, J., Szentgyörgyi, H., Vaissière, B.E., Vaitis, M., Woyciechowski, M., Bazos, I., Biesmeijer, J.C., Bommarco, R., Kunin, W.E., Tscheulin, T., Lamborn, E., Petanidou, T. 2011. Assessing bee species richness in two Mediterranean communities: Importance of habitat type and sampling techniques. Ecological Research, 26:969-983. pdf

Tscheulin, T., and T. Petanidou. 2011. Does spatial population structure affect seed set in pollen-limited Thymus capitatus? Apidologie, 1:67-77. pdf

Schweiger, O., J. Biesmeijer, R. Bommarco, T. Hickler, P. E. Hulme, S. Klotz, I. Kühn, M. Moora, A. Nielsen, R. Ohlemüller, T. Petanidou, S. G. Potts, P. Pysek, J. C. Stout, M. T. Sykes, T. Tscheulin, M. Vila, G.-R. Walther, C. Westphal, M. Winter, M. Zobel, and J. Settele. 2010. Multiple stressors on biotic interactions: how climate change and alien species interact to affect pollination. Biological Reviews 85: 777–795. pdf

Dauber, J., J. Biesmeijer, D. Gabriel, W. E. Kunin, E. Lamborn, B. Meyer, A. Nielsen, S. G. Potts, V. Sõber, J. Settele, I. Steffan-Dewenter, J. Stout, T. Teder, T. Tscheulin, D. Vivarelli, and T. Petanidou. 2010. Effects of patch size and density on flower visitation and seed set of wild plants: a pan-European approach. Journal of Ecology 98:188-196. pdf

Vila, M., I. Bartomeus, A. C. Dietzsch, T. Petanidou, I. Steffan-Dewenter, J. C. Stout, and T. Tscheulin. 2009. Invasive plant integration into native plant-pollinator networks across Europe. Proceedings of the Royal Society B-Biological Sciences 276:3887-3893. pdf

Tscheulin, T., T. Petanidou, S. G. Potts, and J. Settele. 2009. The impact of Solanum elaeagnifolium, an invasive plant in the Mediterranean, on the flower visitation and seed set of the native co-flowering species Glaucium flavum. Plant Ecology 205:77-85. pdf

Woodcock, B. A., S. G. Potts, T. Tscheulin, E. Pilgrim, A. J. Ramsey, J. Harrison-Cripps, V. K. Brown, and J. R. Tallowin. 2009. Responses of invertebrate trophic level, feeding guild and body size to the management of improved grassland field margins. Journal of Applied Ecology 46:920-929. pdf

Potts, S. G., B. A. Woodcock, S. P. M. Roberts, T. Tscheulin, E. Pilgrim, V. K. Brown, and J. R. Tallowin. 2009. Enhancing pollinator biodiversity in intensive grasslands. Journal of Applied Ecology 46:369-379. pdf

Tscheulin, T., T. Petanidou, and J. Settele. 2009. Invasive weed facilitates incidence of Colorado potato beetle on potato crop. International Journal of Pest Management 55:165-173. pdf

Carré, G., P. Roche, R. Chifflet, N. Morison, R. Bommarco, J. Harrison-Cripps, K. Krewenka, S. G. Potts, S. P. M. Roberts, G. Rodet, J. Settele, I. Steffan-Dewenter, H. Szentgyörgyi, T. Tscheulin, C. Westphal, M. Woyciechowski, and B. E. Vaissière. 2009. Landscape context and habitat type as drivers of bee diversity in European annual crops. Agriculture, Ecosystems and Environment 133:40-47. pdf

Westphal, C., R. Bommarco, G. Carré, E. Lamborn, N. Morison, T. Petanidou, S. G. Potts, S. P. M. Roberts, H. Szentgyörgyi, T. Tscheulin, B. E. Vaissière, M. Woyciechowski, J. C. Biesmeijer, W. E. Kunin, J. Settele, and I. Steffan-Dewenter. 2008. Measuring bee biodiversity in different European habitats and biogeographical regions. Ecological Monographs 78:653-671. pdf

Crémieux, L., A. Bischoff, M. Šmilauerová, C. S. Lawson, S. R. Mortimer, J. Doležal, V. Lanta, A. R. Edwards, A. J. Brook, T. Tscheulin, M. Macel, J. Lepš, H. Müller-Schärer, and T. Steinger. 2008. Resistance of local and foreign plant populations to antagonists across a continental scale. New Phytologist 180: 524-533. pdf

Woodcock, B. A., D. B. Westbury, T. Tscheulin, J. Harrison-Cripps, S. J. Harris, A. J. Ramsey, V. K. Brown, and S. G. Potts. 2008. Effects of seed mixture and management on beetle assemblages of arable field margins. Agriculture, Ecosystems and Environment 125:246-254. pdf

Woodcock, B. A., S. G. Potts, E. Pilgrim, A. J. Ramsay, T. Tscheulin, A. Parkinson, R. E. N. Smith, A. L. Gundrey, V. K. Brown, and J. R. Tallowin. 2007. The potential of grass field margin management for enhancing beetle diversity in intensive livestock farms. Journal of Applied Ecology 44:60-69. pdf

Werner, M., T. Tscheulin, T. Speck, D. Zissler, and K. Peschke. 2002. Ultrastructure and motility pattern of the spermatozoa of Aleochara curtula (Coleoptera, Staphylinidae). Arthropod Structure & Development 31:243-254. pdf